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I. Bacteria
a. __major groups of Bacteria, ranging from __ that __to __ such as cyanobacteria and purple and green bacteria
II. Cyanobacteria: Important from Ecological and Evolutionary Views
a. Partake in the global __ and __ and represent one of the major evolutionary lines of Bacteria
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- extreme thermophilic chemosynthetic autotrophs
- oxidize gaseous hydrogen or reduce sulfur compounds
- photosynthetic autotrophs
- carbon and nitrogen cycles
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a. Photosynthetic cyanobacteria have __, together with __and accessory pigments called __(two kinds)
i. What are they?
b. Cells of cyanobacteria have several __, __to each other
i. These membranes are __ that resemble those found in chloroplasts—and, in fact chloroplasts correspond in size to an entire cyanobacterial cell
1. The main storage product of cyanobacteria is __
- chlorophyll a
- carotenoids
- phycobilins
- Phyocoerythrin: red Phycocyanin: blue
- membrane layers
- parallel
- photosynthetic thylakoids
- glycogen
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a. Many cyanobacteria produce a __that bidns groups of cells/ filaments together
i. Colors of the sheaths in different species __
1. “blue-green algae” are not __and are about __
b. They may often form __and grow in large amsses 1 m or more in length
i. Some are __, some have __, and few have __
- mucilaginous envelope (sheath; deeply pigmented)
- vary (are light gold, yellow, brown, etc. )
- algae
- half blue-green
- filaments
- unicellular
- branched filaments
- plates/ irregular colonies
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1. Division--> subunits separate to __
a. Cells of cyanobacteria are joined only by __ or __, so that each cell leads an __life
ii. Some filamentous cyanobacteria are __, doing what?
1. Some segments called __break off from a cy. Colony and glide away from their parent colony at rapid rates
- form new colonies
- their walls or by mucilaginous sheaths
- independent
- motile
- gliding and rotating around the longitudinal axis
- hormogonia
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a. This movement may be connected with the __ through small pores in the cell wall, together with the production of __ in one of the surface layers of the wall
i. Some cyanobacteria exhibit __
- extrusion of mucilage
- contractile waves
- intermittent jerky movements
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b. Where They Live
i. Despite the ‘7500’ species, there may be as few as __
ii. Like other bacteria, they sometimes grow under very __ (hot springs, arctics, etc./ not in acidic places)
iii. Layered chalk deposits called __ (2.7 billion years) are produced when colonies of cyanobacteria bind __
- 200 distinct, free-living nonsymbiotic species
- inhospitable conditions
- stromatolites
- calcium-rich sediments
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1. Today, they are in only a few places
a. Yet, their presence indicates what?
i. Most ancient ones were probably made by __
- that such environments were prevalent in the past, when cyanobacteria played the decisive role in elevating the level of free oxygen in the atmosphere of the early Earth
- purple and green bacteria
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i. Many marine cyanobacteria occur in __, like coralline algae and the shells of mollusks
ii. Some freshwater species of cy. often deposit thick layers of __in their colonies.
- limestone or lime-rich substrates
- lime
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a. Cyanobacterial Formation of Gas Vesicles, Heterocysts, and Akinetes
i. The cells of cy. living in freshwater or marine habitats—especially plankton (on the surface)—commonly contain bright, irregularly shaped structures called __, which do what, thus allowing what? .
- gas vesicles
- provide and regulate the buoyancy of the organisms, thus allowing them to float at certain levels in the water
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1. When several cy. can’t regulate __ properly (maybe due to fluctuating temps), they may __ and __
a. Some cy. that form __secrete __ that are toxic to other organisms and cause death
ii. Many genera of cy. can __[which is available for biological reactions])
- gas vesicles
- float to the surface of the water
- form visible masses called blooms
- blooms
- chemical substances
- fix nitrogen (nitrogenà ammonium
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i. In filamentous cy., the __ often occurs with __, which are specialized, enlarged cells
1. Surrounded by what?
2. Within a __, the cell’s internal membranes are reorganized into a __
- nitrogen fixation
- heterocysts
- thickened cell walls containing large amounts of glycolipid, which impedes diffusion of oxygen in to the cell
- heterocyst
- concentric or reticulate pattern
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a. Heterocysts are low in __, and they lack __, so the __ that occurs in these cells doesn’t result in the __
i. The oxygen that is present is either rapidly __, a by-product of nitrogen fixation, or __
1. __, the enzyme that catalyzes the nitrogen-fixing reactions, is sensitive to the presence of oxygen, and nitrogen fixation is __
- phycobilins
- Photosystem II
- cyclic photophosphorylation
- evolution of oxygen
- reduced by hydrogen
- expelled through the wall of the heterocyst
- Nitrogenase
- anaerobic
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a. __have small __connections—__—with adjacent vegetative cells
2. The products of __ are transported through the __from the __to the vegetative cells, and the products of photosynthesis move in the __ through these same connections, from the __ to the __.
- Heterocysts
- plasmodesmatal
- microplasmodesmata
- nitrogen fixation
- microplasmodesmata
- heterocyst
- opposite direction
- vegetative cells to the heterocyst
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1. Among cy. that fix nitrogen are free-living species such as __, which lives in certain __
a. They account for ¼ of the total __ there
2. __cy. are also important in __
a. In Asia, they helped with growing rice without fertilizers
i. Members of the genus Anabaena often occur in association with the small, floating water fern Azolla, which forms masses on __
- Trichodesmium
- tropical oceans
- nitrogen fixed
- Symbiotic
- nitrogen fixation
- rice paddles
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i. Cy. occur as __within the bodies of a vast number of species: amoebas, sponges, etc. and also act photosynthetically in lichens
1. Some __cy. lack a __, in which case they function as __
2. __ cy. divides at the same time as its host cell by a process similar to __
ii. In addition to __, some cy. form resistant spores called __, which are enlarged cells surrounded by thickened envelopes
1. They are resistant to heat and drought and thus allow the cyanobacterium to survive during unfavorable periods
- symbionts
- symbiotic
- cell wall
- chloroplasts
- Symbiotic
- chloroplast division
- heterocysts
- akinetes
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a. Prochlorophytes contain Chlorophylls a and b and Carotenoids
i. The __are a group of photosynthetic bacteria with __ and __, but no __.
1. Only three discovered: What are they?
- prochlorophyes
- chlorophylls a and b, and carotenoids, but no phycobilins
- prochloron
- prochorococcs
- prochlorothrix
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Prochloron
- a. found only along tropical seashores symbiotically within colonial sea squirts
- i. Spherical cells and extensive thylakoid system
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Prochlorococcus
smallest photosynthetic organism with smallest genome of any photosynthetic cell and most numerous photosynthetic organism on Earth; found in mineral-poor oceans and from surface down to euphotic zone (zone where enough light penetrates for photosynthesis to occur
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Prochlorothrix
a. filamentous and found growing in shallow lakes in Netherlands
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a. Purple and green Bacteria and Photosynthesis
i. Represent second major group of __after cy.
1. The overall __ and the __ used by these bacteria differ from __ and __.
- photosynthetic bacteria
- photosynthetic process
- photosynthetic pigments
- cy. and prochlorophytes
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What is the difference between PnG bacteria and Prochlorophytes and Cy.
- a. Whereas cy. and prochlorophytes produce oxygen during photosynthesis,, the ‘p & g’ bacateria don’t
- i. They grow in light only under anaerobic conditions because pigment synthesis is represented by oxygen
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Differences in photosynthesis
cy: employ chlorophyll a and two PS's in photosynthesis
- Prochlorophytes have chlorophylls a and b and two photosystems
- P and G bacteria: use several different types of bacteriochlorophyll, which differ in certain respects from chlorophyll, and have only one photosystem
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Purple and green bacteria
a. The __are ancestors of individual __—__
i. Unlike p & g bacteria, photosynthetic autotrophs (plants and algae), as well as cy. and prochlorophytes, have __
- photosystems
- Photosystem II and Photosystem I
- both photosystems
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i. Colors characteristic of photosynthetic bacteria are associated with the presence of __ that function in photosynthesis
1. In two groups of purple bacteria= __
2. In cy., the pigments are __, which aren’t in p & g bacteria
ii. Purple and green bacteria are subdivided into those that __
1. __plays same role in photosynthesis that water does
- accessory pigments
- yellow and red carotenoids
- red and blue phycobilins
- use mostly sulfur compounds as electron donors and those that do not
- Sulfur
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i. Purple nonsulfur and green nonsulfur bacteria, which are able to use __ at only low levels, also use organic compounds as __
1. These compounds are fatty acids, alcohols, etc.
- hydrogen sulfide
- electron donors
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1. Because of their requirement for __or a similar substrate, the purple sulfur and green sulfur bacteria can grow only in habitats containing large amounts of __, recognizable by the sulfurous odor
a. In these bacteria, __ may accumulate as deposits within the cell
H2S
decaying organic material
elemental sulfur
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