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H. Heidelbergensis
key fossil sites
Africa
- Africa
- Bodo d' Ar, Ethiopia; 600 kya
- ----thick cranial vault (skull), keeling 1250cc
- ---assoc with acheulean tools
- Broken hill mine, Kabwe, Zimbia; 125 kya
- -----heavy brow, rounded cranium, 1280 cc, prognathic face
- ----many cavities sugg, change in diet
- ---indiviuals found with injuries that show they continue to live on->group support
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H. Heidelbergensis key fossil sites
Greece
- Petralona, Greece 160-240 kya
- -- double arched brow, broad nasal opening, thick cranial bones
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H. Heidelbergensis key fossil sites
Germany
- Steinheim, Germany, 250 kya
- - flat face (canine fossa/ deep root tooth), suprainiac fossa (flat under bun)(neandertal-like), 1100cc
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H. Heidelbergensis key fossil sites
France
- Arago, France, 400 kya
- - 23 individuals, heavy brow, no keeling, 1166cc
- -argo xx1: 1166cc, thick brow, post orbital constriction(pinch by ears), no keel, assoc. with acheulean tools
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H. Heidelbergensis key fossil sites
Europe
- Europe
- 1.Atapuerca, spain-Gran Dolina, 900 kya (aka. h. antecessor)
- --juvenile, small brain (1000cc), double arched brow, large teeth
- 2. sima de los huesos, 300+ kya
- --small brain (1125cc), projecting mid-face (neandertal-like)
- -32 individuals of all ages
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H. Heidelbergensis key fossil sites
Asia
- Asia
- Dali, china; 200 kya
- ---thick cranial vault, keeling, flat face, 1120cc
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h. heidelbergensis
morphology
- -retains cranial robusticity - seen in h.erectus
- -higher, more rounder cranium and a large brain than h. erectus (1200cc)
- -mosaic of traits (e.g keeling(sagittal keeling: h. erectus), canine fossa)
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H. heidelbergensis
Behavior?
Technology, hunting, fire use, mortuary practices
- technology
- Acheulean and wooden tools
- hunting
- -Schoningen, Germany- wood spears, flake tools and butchered horses
- -boxgrove, England- hand axes, butchered rhinos and horse scapula with puncture wound
- fire use
- Terra amata, france
- mortuary practices
- -Bodo- cut marks on face
- -Atapuerca, spain- sima de los huesos, cut marks on gran dolina materials
- -steinheim enlarged foramen magnum for brain extraction
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Neanderthals fossil site
Iraq, Israel, uzbekistan
- Shanidar, Iraq 60 kya
- - 9 individuals, group burial
- Mt. carmel, Israel
- - amud, slender brow, incipient chin, large mastoid
- - kebara 60kya and tabun 110 kya
- Teshik- tash, uzbekistan
- - 9 yr old boy, eastern most distribution
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Shanidar 1
- Neanderthal
- 40 year old male
- healed fractures
- thought to be blind because of damage to optic nerve
- withered arm
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Neanderthals fossil site
France
- La chapelle-aux-saints, france 50 kya
- - arthritic male, "brutish, bent kneed, and not fully erect"
- la ferrassie, france ~70 kya
- - 6 graves "classic Neanderthals"
- St. Cesaire, France; 36 kya
- - Gracile skull, narrow nasal opening, incipient chin
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Neanderthals
fossil site
Croatia
- Krapina, Croatia 130 kya
- - 80 individuals, animal bines and tools, intermediate morphology (individuals more lightly built)
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Classic neanderthal
- receding forehead
- long, low vault
- prominent, double arched brow ridge
- projecting midface
- backward swept cheek bones
- weakly developed chin
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H. Neanderthals
Apomorphic (unique ft)
- large double arched brow ridge
- projecting mid face (retro molar space)
- suprainiac fossa(flat under bun)
- occipital bun
- large cranial capacity, avg. ~1520 (larger than modern humans)
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h. Neanderthals
Symplesiomorphic (ancestral ft)
- small mastoid process
- flat robust skull with a low forehead
- mandible lacks chin
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Neanderthals
morphology
- -long, low cranial vault and large brain (1520cc)
- - occipital bun and suprainiac fossa (elliptic depression on occiput--flat back head)
- -robust double arched supraorbital torus
- - Taurodont molars (fused roots) and shovel shaped incisors- designed for heavy wear
- -robust post cranial skeleton (increased musculature), barrel-shaped chest and shorter distal elements (cold adapted body type)
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Neanderthals
behavior
- Technology
- - Mousterian tool industry (levallois tech) and hafted tools
- - teeth as tools- heavy wear on front teeth
- - chattelperonian tool industry (aft 40 kya)
- ----saint cesaire and arcy-sur-cure, france
- hunting and fire use
- -Kebara- prime adult gazelle and deer, many burned bones, midden
- -high rates of tramatic injuries, esp. head and neck
- mortuary practices
- -post-mortem processing- e.g kapina (broken bones, cut marks, and burning)
- -burial and grave goods- e.g shanidar (flint tools), teshik-tash (ibex horns)
- rituals/ symbolic behavior
- -little or no art (arcy-sur-cure caves in france)
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H.ss fossil sites
europe
- Europe
- 1. Cro-Magnon, france 30kya-fully modern anatomy
- 2. abrigo do lagar velho, Portugal 24.5 kya
- - lapedo child, 4 yrs old, pierced shell necklace and red ochre, mosaic morphology (admixture)
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h. ss fossil site
Asia
- Asia
- 1. mt. carmel, Israel : qafzeh and skhul ~90kya
- - reduced brow, incipient chin
- 2.Dali, china 200 kya
- - also classified as h. heidelbergensis
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H. sapiens sapiens
fossil sites
Africa
- Africa
- 1.omo, Kabish, Ethiopia 198 kya
- -omo 1 (high brow and chin) is > modern than omo 2 (occipital torus)
- 2. Herto, middle awash, Ethiopia 160 kya
- - high cranium, moderate brow, distinct canine fossa, 1450 cc
- 3. Klasies river mouth, south Africa 90 kya
- - presence of chin variable
- -use of aquatic resources
- -hearth
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H. sapiens sapiens
morphology
- -forehead is rounded and vertical
- -canine fossa (deep rooted teeth)
- -pyramidal mastoid process
- - generally gracile jaw and teeth and a distinct chin
- -mean cranial capacity ca. 1400cc
- - limb bones are relatively slender
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h. sapiens sapiens
behavior
MSA tools before ~40 kya (e.g skhul and qafzeh)
- the human revolution
- -greater use of aquatic resources
- -more varieties of stone tools (microliths and blades)
- - use of bone, antler and ivory to make tools
- -stone tool materials transported over long distances
- -burials more complex with grave goods
- -artwork common
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oldowan
acheulean
mousterian
aurignacian
- oldowan
- earliest evidence from h. habilis; disc in olduvai gorge, Tanzania- stone cores with flakes removed from part of the surface, edges made sharp
- acheulean
- disc in France; stone hand axe; pear shaped; used for skinning, digging, butchery, and cutting wood and plants. tools are found far and wide with a production rate that is longer than any other tool
- mousterian
- flint tools, assoc with neadtherals. mid paleolithic, disc in France
aurignacian- made with push blade tech. upper paleolithic, assoc with modern humans, disc in Europe. created blades not flakes, carved bones.
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multiregional ?
- regional contiguiety??
- humans have evolved in multiple regions
- "we all been on species since the beginning"
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replacement or African eve hyp??
when moderns evolved in Africa they moved and replaced older hominins
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assimulation
small pop. assimulated to large pop
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H.ss
canine falsa
- canine falsa
- depression where canines are pressed down under cheek bones
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H. Neanderthals
retro molar space
- retro molar space
- space behind molars
- -no chin
- -no forehead
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Shanidar
teeth as tools
- wear in teeth (none close together)
- hides? skins?
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3 kinds of data
- morphology- mosaic morphology
- archaeology- Neanderthal art and upper paleolithic technology
- genetics- some admixtures
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H. heidelbergensis
ancestral to Neanderthals
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h. antcessor
splits from Neanderthals and early h. ss
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Neanderthal life expectancy
30yrs
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Neanderthals hunting type
close range
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Neanderthal tools technique
- -used precise strikes to make tools (unique)
- - pitch extraction; used as glue to hold spear head to shaft.
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neaderthals and h.ss interbreed?
- 1-4% similar
- Neaderthals genetically close to Europeans and asians
- hybrids in MH pop and conti breeding that's why neaderthal genes continues to be seen in our genome
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h. heidelbergensis
Primitive and derived traits
- primitive
- prognathic face
- thick cranial bones
- large brow ridge
- mandible lacks chin
- derived
- larger brain than h. erectus
- higher rounder cranium than h. erectus
- reduced tooth size
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H. floresiensis
- Disc. 2004
- 18 kya
- 3ft tall
- 417 cc
- no chin
- prominent brow
- femur with long neck
- small teeth
- narrow nose
- thin vault
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Neanderthal post cranial skeleton
- robust post cranial skeleton (increased musculature)
- large joint surface areas
- barrel shaped chest
- short distal elements
- longer superior pubic ramus
- -->larger birth cannel
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Denisovan
- Altai mt, siberia
- one pinky bone
- 2 robust molars
- additional toe bone found with neaderthal DNA
- green stone bracelet found
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